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4 HARVARD UNIVERSITY Library of the Museum of Comparative Zoology 1993VOL 35 MALACOLOGIA MalacologyInternational Journal of MalacologiadeRevista Internacional MalacologieJournal International de ZeitschriftMalakologischeInternationale t»"^ MCZ LIBRARY NO. 1 1993VOL 35, JUL 2 1 1993 HARVARD UNIVERSITY MALACOLOGIA Malacologynternational Journal of MalacologiaRevista Internacional de MalacologieJournal International de ZeitschriftInternationale Malakologische MALACOLOGIA Editor-in-Chief: GEORGE M. DAVIS Editorial and Subscription Offices: Department of Malacology The Academy of Natural Sciences of Philadelphia 1900 Benjamin Franklin Parkway Philadelphia, Pennsylvania 19103-1195, U.S.A. Co-Editors: EUGENE COAN CAROLJONES California Academy of Sciences Denver, CO San Francisco, CA Assistant Managing Editor: CARYL HESTERMAN Associate Editors: JOHN B. BURCH ANNE GISMANN University of Michigan Maadi Ann Arbor Egypt MALACOLOGIA is published by the INSTITUTE OF MALACOLOGY, the Sponsor Members of which (also serving as editors) are: KENNETH J. BOSS JAMES NYBAKKEN Comparative Zoology Moss Landing Manne LaboratoryMuseum of CaliforniaCambridge, Massachusetts JOHN BURCH, President CLYDE F. E. ROPER Smithsonian Institution MELBOURNE R. CARRIKER Washington, D.C. University of Delaware, Lewes W. D. RUSSELL-HUNTER GEORGE M. DAVIS Syracuse University, New York Secretary and Treasurer SHI-KUEI WU CAROLE S. HICKMAN University of Colorado Museum, Boulder University of California, Berkeley President-Elect Participating Members EDMUND GITTENBERGER JACKIE L VAN GOETHEM Secretary, UNITAS MALACOLOGICA Treasurer, UNITAS MALACOLOGICA Rijksmuseum van Natuurlijke Koninklijk Belgisch Instituut Historie voor Natuurwetenschappen Leiden, Netherlands Brüssel, Belgium Emeritus Members Emérita ROBERT ROBERTSONJ. FRANCIS ALLEN, Environmental Protection Agency The Academy of Natural Sciences Washington, D.C. Philadelphia, Pennsylvania ELMER G. BERRY, NORMAN F. SOHL Germantown, Maryland U.S. Geological Survey Reston, Virginia Copyright 1 by the Institute of Malacology© 993